Ok here's my understanding, as for 4WH or 4WL has nothing to do with the hubs that is just a gear change in the transfer case. As for the auto hubs if you are going in reverse it is possible to un-engage a wheel depending which wheel has the most traction and for driving forward is I believe the opposite. Manually locking of the hubs locks them to the axel but then unless you have a limited slip or a locking diff you are still spinning the tire that has the least traction. Like I said, this is my understanding, I could be wrong.