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The reason for keeping the diff 1/4" lower than the hole is so you don't flood the rear hubs with diff fluid. If your hubs have old seals they will probably start leaking if you overfill the diff. It is not hard to change the seals though.
My tunes haven't effected the highway milage, at least it is not noticable. The only things that would really effect highway milage is gearing, trained foot, weight, drag, and perhaps installing an intercooler.
2001 Ford E350 7.3 Extended Cargo Van
BTS Transmission, PHP Minotaur package, 4" exhaust/downpipe, Honey Badger jr., FRX, HPX, AIH delete, Fuel bowl mod, Hypermax Boost/Pyro gauge, 4.10 rear.